Protologue Description: Larnax andersonii N. W. Sawver, sp. nov. TYPE: Ecuador. Napo: km 25 of Holl?n-Loreto road, finca entrance next to bridge over a quebrada in secondary pluvial forest, 950 m, 00°40'S, 7 40'W, 1 Julv 1995, Sawyer & Tirado 714 (holotype, MO: isotypes, CONN, US). Figure 1.

Inter quaterni species, ceteris generis fructu invuluto Iaxe calyce trichomatibus longis et interdum ramosis, ramis secundariis geniculatis, foliis geminatis disparibus, floribus, solitariis et distalibus in ramis, corollis luteolis aut purpuratis, antheris apiculatis sed recedens ab isdem.

Suffrutescent perennial, 0.5-2 m tall. Stems branched, secondary branches geniculate. Stems and leaves villous-sericeous, covered with long, multicellular simple or occasionally branched trichomes. Leaves membranous, entire, paired, unequal, the larger (major) elliptic, 4.5-11 cm long, 1.6-3.9 cm wide with 5-8 secondary veins, the smaller (minor) elliptic-ovate-ovate, 0.3-2.5 cm long, 0.6-1.9 cm wide. Leaves strigose-sericeous above, villose-sericeous below especially along veins, bifurcate hairs occurring, occasionally along leaf margin and along veins; apex acuminate in major leave, acute-mucronate in minor leaves base oblique-equal, atttenuate in major leaves, sometimes attenuate-truncate in minor leaves. Petiole sericeous, 3-10 mm long on major leaves. 0.5-3 on long on minor leaves. Inflorescence distal on the branch, axillary, typically solitary, occasionally paired. Flowers pendent on sericeous pedicels 3-5 mm long. Flowering calyx villous-sericeous, green, 1.5-2 mm long, 2.5-3 mm wide, margin ranging from broadly pyramidal at the 5 major veins to 5-lobed, lobes acute, <1 mm long. Corolla campanulate, 6-8 mm long, tube 2-3 mm long, lobes of the limb membranous. narrowly triangular, sparsely villous externally, densely tomentose along margin and at apex, bifurcate hairs occurring rarely externally and along margin, glabrous within. In shade plants, corolla pale yellow-cream-green; in sun plants, corolla violet with cream base. Stamens included. Filaments glabrous, filamentose, the free portion 1.6-2 mm long, adnate to the basal third of the corolla and there broadened and thickened into a stamen petalum with projections at the shoulders, separated from each other by the corolla vasculature, fused portion 0.9-1.8 mm long. Anthers white-pink, ovate with long apiculum, occurring in two size arrays of three large and two small, 1.3-1.6 mm long, 0.75 mm wide, slightly dorsifixed, base sagittate. Ovary glabrous, ovate. 1 mm long, less than 1 mm wide, ringed basally by a greenish yellow nectary. Style glabrous, 2-4 mm long, extending beyond anthers late in anthesis; stigma clavate-subbilobate. green-purple. Fruiting pedicels sericeous, 5 mm long. Fruiting calyx accrescent, loosely enveloping the fruit, open at the apex, green with dark green veins, villous, membranous, pvriform, 1.2-1.3 cm long, 0.8-1 cm wide, with scattered, two-celled glandular hairs within; lobes distinct. triangular, acute, unequal in length, to 2 mm long. Fruit an orange, fleshy berry containing 6080 reniform, faveolate seeds to 2.8 mm diam.
A suite of characteristics are diagnostic for this species. They include the geniculate younger stems, a character present in many species of this group, and the distinctly unequal-geminate leaves on younger branches; the usually solitary flowers occurring distally on branches: the apiculate anthers also found in several species; and the long, sometimes bifurcate, multicellular hairs also found in L. subtriflora and L. suffruticosa Sawyer, unpublished.

Paratvpes. ECUADOR. Napo: Archidona, faldas al sur del Volcán Sumacu, Carretera Hollin-Loreto, km 31, Comuna . 1200 m. 00°43'S, 77°36'W, 8-17 Jan. 1989. Alvarado 222 (NY); new toad to Loreto (56 km S of Biezal), 28 km E of junction with Baeza-Tena Road. 1100 m. O0°50'S, 77°33'W 21 Dec. 1988. Hammel & Wilder 17263 (NY); El Chaco Canton, Proyecto Hidroeléctrico Coca, Punto ST3. márgen derecha del Río Quìjos, 1500m, 00°11'S, 77°39'W, 3-5 Oct. 1990. Palacios 5805, (QGNE); El Chaco Canton, Proyecto Hidroeléctrico Coca, Punto ST4, Márgen derecha del Río Quijos, ca 10 km al sur de Beventador, 1450 m 00°08'S, 77°30'W, 6-10 Oct. 1990. Palacios 6059 (QCNE) Pastaza: Hacienda San Antonia del Barón von Humboldt, 2 km al NE de Mera,1300 m, 01°27'S, 78°06'W, 27 Feb.-I9 Mar. 1985, Baker, Neill, Palacios & Zaruma, 5662 (MO, NY, QAME); along road from Puyo to Macas, ca. 33 km S of Puyo, 24.9 km S of Veracruz, 16 km S of Escuela Fiscal Cotopaxi, disturbed primary rainforest, 900 m, 01°38'S, 77°52'W, 3 May 1984. Croat 58946 (NY).

Name derivation: The specific epithet honor Gregory J. Anderson, whose dedicated and exhaustive research into the evolutionary and reproductive biology of the genus Solarium sect. Basarthrum, notably his groundbreaking work on cryptic dioecv, warrants recognition. His exemplary, broad-scaled investigation, ranging in contest from experimental research to applied systematic studies- including studies of pollen and hair types, have provided both inspiration and guidance for his students and colleagues.




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